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Landfowl

Landfowl usually refers to birds that mainly move on the ground, forage and breed on land) A more detailed introduction and classification overview. It should be noted that "landfowl" is not a strict monophyletic group, but an "eco-morphological" collection formed based on ecological and morphological characteristics (such as feeding, running or perching on the ground). Modern bird phylogenetic studies show that these ground-dwelling birds are scattered in multiple different orders and have no single common ancestor. Landfowl is one of the six major ecological groups of birds. Landfowl refers to birds in the Galliformes and Columbidae in the class Aves. There are 52 species of birds in these two orders in Yunnan. These birds mainly live on land, so they are called landfowl. Grouse, pheasants, peacocks, etc. all belong to this category. Most landfowl are resident birds, and a small number are migratory birds.

I. Introduction to Terrestrial Birds

1. Definition and Key Traits

  • Primarily Ground-Dwelling: Most terrestrial birds spend significant time foraging, nesting, or resting on the ground. Some (like ostriches and emus) have entirely lost the ability to fly, while others (e.g., many galliform birds) can fly short distances but generally stay on land.

  • Body and Limb Adaptations: They often have strong leg musculature for running, scratching, or digging in the soil. Certain groups specialize in endurance running (e.g., ostriches), while others rely on short, explosive flights for escape (e.g., pheasants).

  • Ecological Diversity: They inhabit a range of environments—tropical savannas, temperate deserts, montane scrublands, and agricultural grasslands. Their diets can be herbivorous, omnivorous, or carnivorous (e.g., eating seeds, insects, small invertebrates).

  • Close Relationship with Humans: Many terrestrial birds have been domesticated (chicken, turkey, guineafowl) or hunted (pheasants, quail). They hold substantial economic and cultural significance.

2. Diversity and Ecological Roles

  • Terrestrial birds span flightless “ratites” (e.g., ostriches, rheas, emus) to ground-adapted fowl that can still fly short distances (e.g., galliform birds like chickens and pheasants).

  • In wild ecosystems, they act as seed dispersers, vegetation controllers, or key links in the food web. In human societies, they provide meat, eggs, and cultural value (e.g., game birds, ornamental species).


II. Evolutionary History

  1. Origins and Radiation

    • Birds emerged from theropod dinosaurs in the Mesozoic era and diversified extensively by the end of the Cretaceous. Some lineages adopted primarily ground-based lifestyles, developing running, scratching, or ground-nesting traits.

    • During the Paleogene and Neogene, geographic isolation and varied habitats spurred further specialization, such as flightless ratites in the Southern Hemisphere and galliforms in the Northern Hemisphere.

  2. Single or Multiple Losses of Flight

    • Large, flightless birds (e.g., ostriches, emus, cassowaries, rheas) exemplify parallel evolution, having independently reduced or lost their flying abilities.

    • Although scattered in different orders/families, they share similar adaptations for terrestrial living (large bodies, powerful legs).

  3. Co-Evolution with Humans

    • Humans have domesticated chickens, turkeys, guineafowl for centuries and hunted many species of pheasants and quail.

    • Several terrestrial birds have significant economic value in agriculture and ecotourism, but habitat loss and hunting threaten some wild populations.


III. Major “Terrestrial Bird” Groups and Classification Table

Below is a list of notable ground-dwelling bird orders and families according to modern avian taxonomy. As “terrestrial birds” are not a monophyletic clade, these orders differ widely in phylogeny but share a ground-based ecological lifestyle.

(A) Ratites and Allies (Flightless “Running Birds”)

1. Ostriches, Rheas, Emus, Cassowaries, Tinamous

These birds typically have lost the ability to fly, are large-bodied, and mostly found in the Southern Hemisphere and Africa, sometimes collectively called “ratites.”

OrderFamilyRepresentative GenusExample SpeciesDistribution & Notes
Struthioniformes (Ostriches)Struthionidae (Ostrich)Struthio (ostrich)Common Ostrich (Struthio camelus)Largest living bird, native to African savannas/deserts; cannot fly, runs at high speed.
Rheiformes (Rheas)Rheidae (Rheas)Rhea (rhea)Greater Rhea (Rhea americana)Found in South American grasslands; flightless, smaller than ostriches.
Casuariiformes (Emus, Cassowaries)Dromaiidae (Emu), Casuariidae (Cassowary)Dromaius (emu), Casuarius (cassowary)Emu (Dromaius novaehollandiae), Southern Cassowary (Casuarius casuarius)Endemic to Australia/New Guinea; large, flightless, known for speed (emus) or powerful kicks (cassowaries).
Tinamiformes (Tinamous)* or included within PaleognathaeTinamidae (Tinamous)Tinamus, CrypturellusGreat Tinamou (Tinamus major), etc.Central & South America; can fly short distances but mostly ground-based. Considered “primitive” among ground birds.

Note: Some classifications treat Tinamous as a separate order (Tinamiformes), while others group them with ratites under Paleognathae, depending on molecular studies.


(B) Galliformes (“Gamefowl”)

Galliform birds are quintessential “land birds”: usually of medium size, with short, rounded wings for bursts of flight but spending most time on the ground.

FamilyRepresentative GeneraExample SpeciesDistribution & Notes
Phasianidae (Pheasants, Chickens, Partridges, Quail, etc.)Gallus (Junglefowl), Phasianus (True pheasants), Coturnix (Old World quail), etc.Domestic Chicken (Gallus gallus domesticus), Ring-necked Pheasant (Phasianus colchicus), Common Quail (Coturnix coturnix), etc.Very diverse, found across Eurasia, Africa, parts of the Americas. Many species are domesticated or important game birds.
Numididae (Guineafowl)Numida (Helmeted Guineafowl), etc.Helmeted Guineafowl (Numida meleagris)Native to Africa, known for “pearl-like” spots on feathers, also domesticated in some regions.
Meleagrididae (Turkeys)Meleagris (turkeys)Wild Turkey (Meleagris gallopavo)Native to North America. Large ground birds, source of domestic turkey; males exhibit colorful wattles and tail fans.
Tetraonidae (Grouse, Ptarmigan)* sometimes included in PhasianidaeTetrao (Capercaillies), Lagopus (Ptarmigan)Western Capercaillie (Tetrao urogallus), Willow Ptarmigan (Lagopus lagopus), etc.Typically in cooler Northern Hemisphere regions or high mountains. Feathers can change seasonally for camouflage.
Odontophoridae (New World Quail)Colinus (Bobwhites), Odontophorus (Wood Quail), etc.Northern Bobwhite (Colinus virginianus), etc.Found in the Americas, resembling Old World quail in body form and ground-based habits.

Note: Family boundaries within Galliformes vary; some sources merge grouse into Phasianidae.


(C) Other Common Ground-Dwelling Birds

Beyond the “ratites” and “gamefowl,” several other orders have predominantly ground-based families or genera. A few key examples:

  1. Otidiformes (Bustards)

    • Otididae (Bustards): e.g., Great Bustard (Otis tarda), African bustards; large, flight-capable but strongly ground-oriented birds in Eurasian and African grasslands.

  2. Gruiformes (Cranes, Rails, and Allies)

    • Rallidae (Rails, Coots): e.g., Common Coot, various rails. Primarily live in wetlands or marshy ground.

    • Gruidae (Cranes): Though cranes migrate via flight, they feed and nest on the ground in marshes or grasslands.

  3. Columbiformes (Pigeons, Doves)

    • Many species are strong fliers, but certain doves and ground-doves spend much time pecking seeds on the ground.

  4. Passeriformes (Perching Birds) partial ground-dwellers

    • Families such as Alaudidae (Larks) and some Motacillidae (Wagtails, Pipits) forage extensively on open ground, though they remain capable of sustained flight.


IV. Summary

  1. Multiple Origins and Convergent Adaptations

    • “Terrestrial birds” do not form a single evolutionary lineage; rather, multiple avian orders independently evolved ground-based lifestyles—strong legs, ground nesting, and foraging.

    • From large flightless ratites to ground-frequenting galliforms and bustards, they illustrate repeated occupation of terrestrial niches.

  2. Ecological and Human Importance

    • Many ground birds serve crucial functions in grassland or desert ecosystems, dispersing seeds, controlling insect populations, etc.

    • Humans have domesticated chickens, turkeys, guineafowl, and more. Hunting and wildlife tourism also focus on species like pheasants and bustards.

  3. Conservation Concerns

    • Habitat loss, agriculture expansion, and overhunting threaten many terrestrial bird populations (e.g., ostriches, cassowaries, great bustards).

    • Species with limited ranges or migratory needs require international collaboration and protected areas for survival.


Through the above classification and introduction, your website users can have a preliminary understanding of the diverse distribution and evolution of the ecological-morphological group "land birds" in the modern bird system. If you need to further understand the morphological characteristics, ecological habits and conservation status of each species, it is recommended to refer to professional ornithology, regional field manuals and the latest molecular systematics research.

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Caloenas nicobarica

Caloenas nicobarica

Caloenas nicobarica,Nicobar Pigeon,Nicobar Dove,Paloma de Nicobar

Features:It is stocky and has a distinctive long bark on its neck

Caloenas nicobarica has two subspecies: Nicobar Pigeon, Nicobar Dove, and Paloma de Nicobar.A highly wandering bird, the named subspecies was recorded in November 2009 from Ponape, Federated States of Micronesia, 1500 km from the closest known residence of the species (in northeastern Melanesia). Tr...

Uropelia campestris

Uropelia campestris

Uropelia campestris,Long-tailed Ground-dove

Features:

Uropelia campestris and Long-tailed Ground-dove are unknown.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1: Red List of Birds 2010.Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's responsibility!...

Oena capensis

Oena capensis

Oena capensis,Namaqua Dove

Features:The vision range can reach 300 degrees

Its scientific name is Oena capensis and its foreign name is Namaqua Dove.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1: Red List of Birds 2010.Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's responsibility!...

Spilopelia senegalensis

Spilopelia senegalensis

Spilopelia senegalensis,

Features:The outer tail feathers are white with distinctive bluish-gray wing spots

The brown turtle dove (Spilopelia senegalensis) has six subspecies and is a pink-brown turtle dove.The turtledove is a resident bird. Usually in pairs or in small groups of 3-4 individuals, but in places where water and food sources are abundant, large groups, even hundreds of individuals, gather. B...

Henicophaps foersteri

Henicophaps foersteri

Henicophaps foersteri,New Britain Bronzewing

Features:

The species is known as Henicophaps foersteri and New Britain Bronzewing.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1:2010 Red List of Birds - Vulnerable (VU).Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's responsibility!...

Henicophaps albifrons

Henicophaps albifrons

Henicophaps albifrons

Features:

White-topped ground pigeons are known as Henicophaps albifrons and their specific habits are unknown.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1: Red List of Birds 2010.Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's respo...

Metriopelia morenoi

Metriopelia morenoi

Metriopelia morenoi,Bare-eyed Ground-dove,Moreno's Ground Dove

Features:

Metriopelia morenoi, Bare-eyed Ground-dove, Moreno' s Ground Dove, whose specific habits are unknown.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1: Red List of Birds 2010.Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's r...

Metriopelia ceciliae

Metriopelia ceciliae

Metriopelia ceciliae,Bare-faced Ground Dove

Features:

The behavior of Bare-faced Ground Dove (Metriopelia ceciliae) is unknown.Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's responsibility!...

Metriopelia aymara

Metriopelia aymara

Metriopelia aymara,Golden-spotted Ground-dove

Features:

Its scientific name is Metriopelia aymara, and its foreign name is Golden-spotted Ground-dove.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1: Red List of Birds 2010.Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's responsibili...

Metriopelia melanoptera

Metriopelia melanoptera

Metriopelia melanoptera,Black-winged Ground-dove

Features:

Metriopelia melanoptera and Black-winged Ground-dove are unknown.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1: Red List of Birds 2010.Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's responsibility!...

Reinwardtoena crassirostris

Reinwardtoena crassirostris

Reinwardtoena crassirostris,Crested Cuckoo-dove

Features:

Its scientific name is Reinwardtoena crassirostris and its foreign name is Crested Cuckoo-dove.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1:2010 Red List of birds - Near Threatened (NT).Protect wild animals and eliminate wild meat.Maintaining ecological balance is ever...

Reinwardtoena browni

Reinwardtoena browni

Reinwardtoena browni,Pied Cuckoo-dove

Features:

The turtle is Reinwardtoena browni, Pied Cuckoo-dove, and its behavior is unknown.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1:2010 Red List of birds - Near Threatened (NT).Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's re...

Reinwardtoena reinwardtsi

Reinwardtoena reinwardtsi

Reinwardtoena reinwardtsi,Great Cuckoo-dove

Features:

Its scientific name is Reinwardtoena reinwardtsi, and its foreign name is Great Cuckoo-dove.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1: Red List of Birds 2010.Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's responsibility...

Claravis godefrida

Claravis godefrida

Claravis godefrida,Purple-winged Ground-dove,Purple-barred Ground-dove,Purple-winged Ground Dove

Features:

Its scientific name is Claravis godefrida, and its foreign names are Purple-winged ground-dove, Purple-barred ground-dove, Purple-winged Ground Dove.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1: Red List of Birds 2010.Protect wild animals and eliminate wild meat.Mainta...

Claravis mondetoura

Claravis mondetoura

Claravis mondetoura,Maroon-chested Ground-dove

Features:

]...

Claravis pretiosa

Claravis pretiosa

Claravis pretiosa,Blue Ground-dove

Features:

Its scientific name is Claravis pretiosa and its foreign name is Blue Ground-dove.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1: Red List of Birds 2010.Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's responsibility!...

Phaps histrionica

Phaps histrionica

Phaps histrionica,Flock Bronzewing

Features:

The scientific name Phaps histrionica, the foreign name Flock Bronzewing, the specific habit is unknown.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1: Red List of Birds 2010.Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's re...

Phaps elegans

Phaps elegans

Phaps elegans

Features:

Copper-winged brush doves are known as Phaps elegans, but their specific habits are unknown.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1: Red List of Birds 2010.Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's responsibility...

Phaps chalcoptera

Phaps chalcoptera

Phaps chalcoptera

Features:

The copper-winged turtle is known as Phaps chalcoptera, but its behavior is unknown.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1: Red List of Birds 2010.Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's responsibility!...

Geopelia striata

Geopelia striata

Geopelia striata,Zebra Dove,Barred Ground Dove

Features:

Geopelia striata, Zebra Dove, Barred Ground Dove, the specific habit is unknown.Listed in the International Union for Conservation of Nature (IUCN) ver 3.1: Red List of Birds 2010.Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's responsibility!...

Geopelia humeralis

Geopelia humeralis

Geopelia humeralis,Bar-shouldered Dove

Features:

Geopelia humeralis (geopelia humeralis) and Bar-shouldered Dove (Geopelia humeralis).Listed in the International Union for Conservation of Nature (IUCN) ver 3.1: Red List of Birds 2010.Protect wild animals and eliminate wild meat.Maintaining ecological balance is everyone's responsibility!...